There are many little "gotchas" with setting up incoming email for MOSS. Just nit picky little things, normally they are all about email infrastructure, SMTP, etc. My advice below is to help you avoid taking your heart medication, it can be easy to set this up, but it can also be a hair pulling, eye rolling, staying up all night and getting no results kind of affair as well.
I promise, this technology does actually work, but if you deviate from the standard build (see list item 3 below for "standard build" as defined by me) you will fall off a cliff unless really understand what you are doing.
I'm not an exchange guru, just a humble BizTalk/Moss guy.
Don't just do this trial and error and play with settings until it works. It won't work. Be deliberate and intentional, setup the standard way and then slowly change things, each time checking to see if the forwarding still works.
This is where I recommend you start, read all this stuff first, before you race off back to your Virtual Machines.
If you try to read from the SMTP queue on the MOSS server, you will get
A critical error occurred while processing the incoming e-mail file C:\Inetpub\mailroot\Queue\NTFS_cab371a601c8166600000011.EML. The error was: The process cannot access the file 'C:\Inetpub\mailroot\Queue\NTFS_cab371a601c8166600000011.EML' because it is being used by another process..
The solution to this is "DON'T READ FROM THE QUEUE". If the emails are in the SMTP queue on your Moss server then your email/smtp settings are probably messed up. (Perhaps you did not take my advice on keeping the FQDN of the MOSS server the same as the @domain name that you are sending mail to?)
______________________
If you try to "drag and drop" from queue to the drop folder manually, you will get
A critical error occurred while processing the incoming e-mail file C:\Inetpub\mailroot\drop\NTFS_db489bee01c8166600000012.EML. The error was: Bad senders or recipients..
Don't tell me, let me guess. You copied the .eml files from the Queue folder to the Drop folder and then saw sharepoint delete them. Same point as above, If the emails are in the SMTP queue on your Moss server then your email/smtp settings are probably messed up. (Perhaps you did not take my advice on keeping the FQDN of the MOSS server the same as the @domain name that you are sending mail to?)
_______________________